Optimus Prime is dead. Or at least, that's what it seems. But if he's down, what chances are left for the Autobots agains the Mighty Megatron? Meanwhile, the time is expiring not only for Verity, Jimmy, Ironhide and Ratchet, but also for Hot Rod who is under heavy fire. He's in charge of the facsimile but Skywarp and Thundercraker won't let him flee that easily.
With the power balance between Autobots and Decepticons lost, the fragile peace on Earth is about to end. Finally, Devastation is upon us.