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On Earth, a mammoth new Decepticon named Trypticon destroys the Ark and heads toward Autobot City. On Cybertron, a Decepticon armada attack while the Autobots' planetary defenses are down. SkyLynx and the Predacons each head to Io to secure the wayward Transformation Cog - but the Quintessons have an ace up their sleeve, planning a betrayal of Galvatron and the immediate destruction of all Transformer live on Cybertron.

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