Josie and the Pussycats are a rock band from Riverdale founded by Josie McCoy and Melody Valentine. They consist of Josie McCoy on guitar and lead vocals, Valerie Brown on bass (although also seen playing tambourine and baritone guitar) and backing vocals, and Melody Valentine on drums and backing vocals. The Pussycat's are managed by Alexander Cabot III with the occasional help of his twin sister Alexandra Cabot whom fancies herself the forth member of the Pussycats.
The girls perform in trademark pussycat costumes but when not performing they wear casual attire.
Although they are in in the same continuity of the rest of the Riverdale residents, more often than not they stick to their own stories, and interact with the other more popular characters less frequently.