Captain America (HYDRA) appears in 87 issues.
Scientist Arnim Zola became one of the first biochemists in history after finding notes and equipment belonging to the offshoot race of humanity, the Deviants. He learned many things from his studies, including the ability to transfer people's essence into clones of their original bodies.
The 13th Baron in the Zemo bloodline. Helmut followed in the footsteps of his father before him, becoming an enemy of Captain America and the Avengers. In what could be his greatest scheme ever, Helmut masqueraded as Citizen V and led the Thunderbolts to public acclaim in order to conquer the world. In the end, he failed only because his cohorts desired to remain heroes and rebelled against him.
Doctor Faustus is a villain mostly conspiring against Captain America. He deems himself "Master of Mens' Minds" for using psychological combat against his enemies.
A strange woman who introduced Hydra to Sarah and Steve Rogers.
Tomi Shishido is a mutant extremist that formed the "Dawn of the White Light" cult. He has served as a high-ranking official of The Hand and HYDRA. Shishido ranks among the most skilled swordsmen of the world.
Kraken was a master manipulator and a founding father of HYDRA. He specialized in helping people become who they were supposed to be. Secretly, he was killed and replaced by Jake Fury.
The mother of Steve Rogers.
Viper is the former terrorist leader of HYDRA and an enemy of Captain America, Nick Fury and Wolverine among many others. While she is not the ruler of Madripoor, she did - or does - own it.
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